The Integrator block in Simulink takes an input signal u(t) and outputs its integral over time, starting from an initial condition. The output y(t) at each step is the sum of the previous output plus the current input value. For example, with input values 1, 2, 3, 4 and initial condition 0, the output progresses as 1, 3, 6, 10. This cumulative sum behavior models how quantities accumulate over time. The initial condition sets the starting point of the integral. If the input is zero, the output remains unchanged from the previous step. This behavior is essential for simulating systems with accumulation or memory.